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Radiated humans or slightly radiated humans who got diped (not enough to kill them and enough for them not to become supoer mutants).
And Jesus came unto thee... oops, sorry...
Ghouls or 'G00ls' if you are l337 are simply humans who have been exposed to radiation and have changed over a long period of time as a result. Super Mutants are the only outcome of being 'dipped', that's why there are no ghouls on the master's army.
...and he said 'love thy neighbour' to which the stranger replied 'well who is my neighbour, stop being so damn vague, 'son of god'" And then Jesus said 'Ah bugger it all'.
Ummm... I think the coons would have been sterile.
I think the ghouls got some FEV and rad. I think they are screwed up because they have genetic damage. Talius, a former Vault 13-dweller says he turned into a ghoul, because the Nightkin got a hold of him. Which might mean that he was dipped, and came out bad. Since he came out bad, they let him go. Of course, i have no idea what the game designers intended, if anything.
I really liked the plot, it was a cool idea. Now if they could only get rid of the bugs...
